有構厲害|植物學家的人類學之旅
有構厲害|植物學家的人類學之旅
2018/09/04
2021/05/24
<i><b></b></i>
展覽頁面呈現形式
縱式
策展者
章璿
鍾國芳
展覽封面
5d5477a3a06fc2212277efb1c47f68aa
展覽資料
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"desc": "<b>構樹是臺灣最常見的植物之一,常為荒廢空地或林地中的優勢種,卻也是最為人視而不見的植物。你能想像路邊牆腳下那棵從不留人眼光的構樹,竟乘載著人類遷移史上最偉大的一段航程嗎?</b><br><br><b>2008年3月,因緣際會下,現中研院生物多樣性研究中心鍾國芳副研究員團隊開始投入構樹的研究,由此串聯起一系列涵蓋植物系統分類學、生物地理學、生物人類學、考古人類學、語言學、人類遺傳學、民族學的研究篇章。「有構厲害」數位系列展覽以此為本,將從不同面向、以三大主題介紹構樹的故事。 </b><br><br><b>生物學如何能回答人類學的提問?<br>1990年代起,科學家們便嘗試以共生物種的親緣地理研究為南島語族拓殖與往返島嶼間的複雜歷史理出頭緒。2015年11月,鍾國芳副研究員團隊的研究成果於國際期刊PNAS上發表,結合植物學、植物分類學、DNA</b><span><b>親緣地理學、演化生物學、語言學、考古學、人類遺傳學、民族學等的跨領域研究</b>,</span><b>以構樹的親緣地理反映了南島語族遷徙的全貌,人類學家的難題,或許可由植物學家來解答。</b>",
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"desc": "<div>由2005年萬那杜官方發行的拉匹達文化郵票,我們得以一探早期太平洋南島語族先民的生活。<br></div><div>拉匹達遺址多在海岸地區,<b>人們群居並形成村落</b>,搭建干欄式房屋或茅草、木屋,<b>使用陶製及石製的器具</b>,也會生火煮食。當時的人們很<b>依賴海洋資源</b>,採貝、網魚,或是射魚維生。<b>農業也可能是主要的生產方式</b>,以種植根部作物和樹木作物為基礎,最重要的是芋頭、山藥、香蕉、麵包樹、構樹等,作為食物之外,植物也可用來製作日常生活用品,像是編織成容器或製作成穿戴的衣物;除此之外,也會<b>飼養豬,狗和雞等牲畜</b>。</div><div>這些拓殖者將人類的住地擴展到了整個太平洋,成功的關鍵就是<b>「地景轉移」</b>,他們會將這樣的生活模式一併攜帶複製到下一個居所。當南島語族人遷徙時,熟悉的工具、用品、食物,以及一些較快繁殖的動植物會一併打包上船,除了能夠應付險峻的海上生活,也讓他們在找到下一個島嶼家園時,可以快速的安身立命。<br><br><br><div><b><i>拉匹達文化</i></b><i>(Lapita culture)為太平洋地區的史前文化,根據目前出土的代表性陶器,這個文化約3千多年前在近大洋洲的俾斯麥群島乍然而現,隨後離開近大洋洲,向萬那杜等地遷徙,進入遠大洋洲,來到斐濟、東加、薩摩亞一帶。</i><i>這些拉匹達人口中說的正是南島語,而</i><i>拉匹達人則被認為是現今美拉尼西亞與波里尼西亞人的祖先。</i></div><div><i>近大洋洲早在數萬年前便有人居住,拉匹達文化與這些早期的文化有相當大的差異,且是這個地區首次出現使用陶器的文化。</i><i>拉匹達文化</i><i>以複雜的紅土印紋陶器與貝殼和黑曜石製成的工具著名,如此成熟的文化不太可能在一夕之間形成,它是從何興起,又是如何傳遞,一直是難解的謎,許多考古學者認為拉匹達文化是南島語族所創,並隨之擴散、進駐太平洋,而許多關於南島語族遷徙途徑的研究亦多從拉匹達文化的考古、人類及遺傳學著手。</i></div><div><i><br></i><a target=\"_blank\" rel=\"nofollow\" href=\"http://lapita.rchss.sinica.edu.tw/web/\">太平洋史前Lapita陶器線上數位資料庫</a><i><br></i></div></div>",
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"desc": "<span>這一系列的假說憑藉的多是語言與考古學上的證據,語言學家白樂思 (Robert\nBlust) 自1970年代開始研究南島語族的文化史與起源,從南島語系的演化支序圖來看,臺灣擁有最高的多樣性,極有可能為南島語的發源地,也因此提出<b>「臺灣原鄉論」</b>的說法。考古學家彼得.貝爾伍德 (Peter Bellwood) 則將考古資料整合語言學的研究成果,認為南島語族的遷徙是由北往南的農業擴張,並進一步提出<b>「出臺灣說」(Out of Taiwan hypothesis)</b><span>。<br></span></span>生物學家賈德.戴蒙 (Jared Diamond) 以此為基礎,於1988年提出了<b>「快車說」(Express train model)</b>,認為南島語族的農民祖先在距今約6000年前經由中國南方遷移到臺灣。有一部分的人群又在距今5500百年前離開臺灣朝南方遷移,距今約5000年前進入印尼,在接下來的2300年內橫越了約1萬公里的海洋,穿過菲律賓、印尼,約3500年前進入近大洋洲的美拉尼西亞島群,並在3200年前越過新幾內亞來到蘇門答臘群島,抵達遠大洋洲玻里尼西亞的西端,約在距今3000年前進入玻里尼西亞,南島語族人持續向東擴張,並同時向南及向北遷徙,在距今約1200年前抵達其分布北界的夏威夷、距今約1200到800年前到達東界的復活節島,最終在約730年前來到最南界的紐西蘭。<br>依照如此遷移的路徑及時間,南島語族在約2000年的時間內一路從臺灣橫跨整個太平洋,<b>中間沒有停頓,也沒有與當地原居民交流</b>,以當時有限的航海能力展現驚人的拓殖速度,就像一部開往玻里尼西亞的特快列車。<br>",
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"desc": "除了從語言、考古、人類遺傳學來探討南島語族的遷徙,1994年人類學家伊莉莎白‧馬諦索史密斯 (Elizabeth Matisoo-Smith) 提出了<b>「共生物種」(commensal species)</b> 的概念。<br>人類生活在自然之中,其對食、衣、住、行、育、樂等需求,以及自身文明、文化的發展無形之間也慢慢改變了那些與生活關係密切的動、植的自然地理分布和遺傳結構。<br><br>南島語族拓殖成功的關鍵為地景轉移,他們遷移時會攜帶整套的<b>「農業包裹」(agriculture package)</b>,打包所有食、衣、文化所需的原料,如豬、雞、老鼠、麵包樹、芋頭、地瓜等。這些動植物並非原來就分布於該地,太平洋地區約有70種植物並非當地的原生種,且本身並不具有長距離涉水移動的能力,皆是藉由南島語族的遷徙而傳播過去的,南島先民的生存與文化傳承產生緊密的共生關係,所以<b>研究這些物種的族群結構,或許便能夠推論影響他們的南島語族人當初是如何活動的。<br></b><b><br></b><br><a target=\"_blank\" rel=\"nofollow\" href=\"http://www.pnas.org/content/101/24/9167\">緬甸小鼠的研究支持 T</a><a target=\"_blank\" rel=\"nofollow\" href=\"http://www.pnas.org/content/101/24/9167\">riple I假說</a><br><a target=\"_blank\" rel=\"nofollow\" href=\"http://science.sciencemag.org/content/323/5913/527\">胃</a><a target=\"_blank\" rel=\"nofollow\" href=\"http://science.sciencemag.org/content/323/5913/527\">幽門桿菌的研究支持出臺灣說</a><br><a target=\"_blank\" rel=\"nofollow\" href=\"http://www.pnas.org/content/104/12/4834\">太平洋豬源自中南半島</a><br><a target=\"_blank\" rel=\"nofollow\" href=\"https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0039171\">雞骨化石顯示南島語族在1千年前</a><a target=\"_blank\" rel=\"nofollow\" href=\"https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0039171\">已橫</a><a target=\"_blank\" rel=\"nofollow\" href=\"https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0039171\">越太平洋</a><br><a target=\"_blank\" rel=\"nofollow\" href=\"https://onlinelibrary.wiley.com/doi/full/10.3732/ajb.91.5.760\">大洋洲麵包樹的起源交雜古代及現代人類的移動</a><br><br>這些關於共生物種的親緣地理研究,揭開了南島語族拓殖與往返島嶼間的複雜歷史,有時能清楚的支持某個論點,有時卻更加點出了南島語族遷徙這個議題的盤根錯節。<br>",
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"desc": "在當地植物學家的陪同下進入人煙罕稀的森林中尋找構樹。",
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"desc": "田野調查時,各種突發狀況都是家常便飯,車子突然拋錨、工具損壞、迷路或甚至人員受傷,一切都須仰賴團隊的在野外的應變能力,也十分感謝當地的學者、合作夥伴、鄉里居民的大力協助。所以每一次的野外採集都是極耗資源的,出發前必定得謹慎規劃,準備萬全。",
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"desc": "斐濟當地居民對於書中描寫關於自己文化的內容深感興趣,聚精會神的閱讀,一旁的孩童則是被書中照片吸引,興致勃勃的討論。",
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"title": "東加_市集中的樹皮布",
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"desc": "現今夏威夷樹皮布文化復甦源於1960年代Malia Solomon女士的推廣。<br>Malia Solomon為夏威夷原住民,1950年代時移民到加州,與丈夫一同經營外燴事業,經營有成並累績一定財富後,Solomon女士一心希望能夠回饋故鄉,於1960年代回到夏威夷,創立第一個文化村。當時比夏博物館人類學家Kenneth Emory博士建議她可以研究在夏威夷已經消失已久的傳統樹皮布文化,Solomon女士便開始積鑽研樹皮布文化與參加各類相關活動。Emory博士在一次研討會上認識了來自日本白石市的和紙工匠遠藤忠雄先生,並討論了樹皮造紙與製布的技術,在Emory博士的引薦下,Solomon女士於1967年親至日本白石市學習和紙工藝。<br>圖中白石和紙展示館工作人員向大家展示Solomon女士寄來的蒐集自太平洋各地的樹皮布。",
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"desc": "向大家展示過去Solomon女士與片倉先生的通信,以及相互交流寄送的各式樹皮布。",
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"desc": "特別向大家展示以和紙製成的和服。",
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"title": "巴布亞新幾內亞",
"desc": "2018年9月,鍾國芳副研究員(右一)與史前館張至善博士(左一)前往巴布亞新幾內亞進行田野調查,與森林研究所所長(PNG Forestry Research Institute)(右二)及當地標本館人員(左二)合影。 ",
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"desc": "史前館張至善博士細心觀察採集構樹。",
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"title": "請輸入頁面標題",
"desc": "當地居民向研究團隊展示村莊中栽植的構樹。",
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"desc": "研究團隊在村民的引導下尋找構樹,當地孩童好奇又熱情的在一旁圍觀嬉鬧。",
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"title": "菲律賓田野調查",
"desc": "鍾國芳副研究員於2018年9月赴菲律賓採集構樹。",
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"title": "菲律賓",
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"desc": "請輸入頁面說明",
"weight": 59
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"title": "構樹",
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},
"title": "請輸入頁面標題",
"desc": "研究團隊更是在臺灣全島都留下了這樣辛苦採集構樹的身影。<br><br>進行田野採集時,條件時常是非常嚴苛的,有時需要去到環境較為落後的地區,光是到達該地便十分困難,還要擔心在野外調查時的安危;有時語言無法相通,又沒有足夠的相關文獻,讓採集毫無頭緒。團隊成員們耗費許多心力規劃、聯絡,竭盡所能的運用可得的資源,在橫跨將近1萬公里的採集範圍中,<b>總計包含了604個構樹樣本</b>。",
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"level": "primary",
"title": "構樹的葉綠體基因組",
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"fileLicence": "著作權利所有",
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},
"title": "分子標記-構樹葉綠體基因組",
"desc": "<span>要分析構樹的親緣地理,首先得在眾多的分子標記中,找出最適合構樹的那個,才能得到最佳的解析度。在選擇不同的基因片段時,需要考慮物種、樣本數、研究尺度以及實驗可行性等各式不同因素,經過多次的測試,終於發現葉綠體中<b> </b><i><b>ndh</b></i><b>F-</b><i><b>rpl</b></i><b>32 </b></span><b>基因間隔區 (intergenic spacer) </b>(綠色框框處)能夠提供足夠的遺傳訊息。",
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"level": "primary",
"title": "構樹葉綠體 ndhF-rpl32 基因單型網狀圖/構樹葉綠體 ndhF-rpl32 基因單倍型網狀圖",
"desc": "各基因單型 (haplotype) 以不同色系表示,並標上編號1-48,跨號中的數字為樣本數,圓圈的大小與樣本數成比例,圓圈越大表示攜有該基因單型的樣本越多,小於10的樣本數未標示。出現在臺灣的基因單型以虛線表示,方形則表示該基因單型為特有基因單型 (unique haplotype)。",
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},
"title": "構樹葉綠體 <i>ndh</i>F-<i>rpl</i>32 基因單型網狀圖",
"desc": "<div>604個構樹樣本中,一共找到了<b>48個基因單型</b>,並將他們之間的關係畫成網狀圖,一個圓圈 (或方形) 即為一個基因單型,用不同顏色區別,並在內標示編號,圓圈的大小與樣本數成正比,實際數字標示於括號內,小於10的樣本數未標示。48個基因單型中,<b>出現於臺灣的有19個</b>,以虛線標出。</div>",
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"level": "primary",
"title": "構樹基因單倍型的地理分布",
"desc": "將不同的基因單型依照分布地區著色;日本:淡橘色,中國:黃色,臺灣:綠色,菲律賓:紫色,印尼:深藍色,近大洋洲:淺藍色,遠大洋洲:橘紅色。\r\n由圖中可以看出,臺灣、中國與中南半島有絕大多數的基因單倍型,為構樹遺傳多樣性中心,極有可能為構樹的原生地。\r\n",
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"title": "構樹的原生地",
"desc": "若將網狀圖依照各基因單型分布的地理區重新上色,可以發現呈現最多顏色為的為<b>黃色(中國)、綠色(臺灣)及深藍色(中南半島)</b>,分別帶有27、19和5個基因單型。基因多樣性較高的區域,最接近族群散佈的起點,由此可以確定,<b>臺灣、中國與中南半島爲構樹遺傳多樣性中心,是構樹的原生地</b>。<br><br><br>",
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"title": "構樹葉綠體 ndhF-rpl32 基因單型分布及網狀圖/構樹葉綠體 ndhF-rpl32 基因單倍型分布及網狀圖",
"desc": "將各基因單型 (haplotype) 以不同色系表示,並標上編號1-48,圖中可見各基因單型於地理上的分布以及網狀圖。\r\n在地圖上以圓餅圖標示不同基因單型於同一地理區所佔的比例,圓餅圖的大小對應該地的樣本數。在網狀圖中各基因單型的編號標示於圓圈內,跨號中的數字為樣本數,圓圈的大小與樣本數成比例,圓圈越大表示攜有該基因單型的樣本越多,小於10的樣本數未標示。出現在臺灣的基因單型以虛線表示,方形則表示該基因單型為特有基因單型 (unique haplotype),僅出現在特定地理區。太平洋地區紅圈內填黑色代表標本樣本。",
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"title": "太平洋構樹如何傳播",
"desc": "進一步將基因單型依其分布繪於地圖上,再對照網狀圖,當中的脈絡就更加分明。<br><br>首先可以發現的是,臺灣雖然面積狹小,分佈在臺灣的構樹卻有非常高的單型多樣性,而且構樹的族群在北部(綠色系)、中南部(紅色系)與東部(藍色系)的單倍型屬於不同演化支序,顏色在地圖上的呈現更可以看出明顯的分群;<b>顯著的親緣地理結構暗示著,構樹雖然是常見植物,但其自然傳播力有一定的局限性,更遑論越洋傳播。<br></b><br><b>出南臺灣</b><br>而與原生地相比,<b>太平洋上的基因多樣性則明顯低落,在蘇拉威西、新幾內亞及遠大洋洲島嶼上,絕大多數的構樹均攜帶紅色編號17的基因單型(cp-17),而除了大洋洲外,cp-17僅分布在臺灣南部</b>。cp-17位於一個臺灣特有的基因單型群的最末端,即網狀圖中紅色系的分支,由遺傳分析判斷,一定是由分布在臺灣南部的橘色cp-9和紫紅色cp-16演變而來。而極低的多樣性同時也是<a target=\"_blank\" rel=\"nofollow\" href=\"https://zh.wikipedia.org/wiki/%E9%81%97%E4%BC%A0%E6%BC%82%E5%8F%98#%E7%A7%8D%E7%BE%A4%E7%93%B6%E9%A2%88\">瓶頸效應(bottleneck effect)</a>與<a target=\"_blank\" rel=\"nofollow\" href=\"https://zh.wikipedia.org/wiki/%E9%81%97%E4%BC%A0%E6%BC%82%E5%8F%98#%E7%A7%8D%E7%BE%A4%E7%93%B6%E9%A2%88\">建立者</a><a target=\"_blank\" rel=\"nofollow\" href=\"https://zh.wikipedia.org/wiki/%E9%81%97%E4%BC%A0%E6%BC%82%E5%8F%98#%E7%A7%8D%E7%BE%A4%E7%93%B6%E9%A2%88\">效應(founder effect)的信號</a>,表示這些遠大洋洲帶有cp-17的構樹最有可能是被帶出臺灣至遠大洋洲的共生物種。cp-17在臺灣與大洋洲間的不連續分布無疑證明了<b>臺灣是「太平洋構樹」的原鄉</b>,如此看來,太平洋構樹的親緣地理與臺灣是南島語族原鄉的<b>「出臺灣說」</b>十分吻合。<br>",
"weight": 64
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"title": "福建至華中及臺灣北部地區的基因單型",
"desc": "福建至華中一帶最常見的單倍型 cp-1亦為北臺灣構樹主要攜帶的基因單型。",
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"title": "出南中國、入北臺灣",
"desc": "<b>北臺灣單型多樣性較低,幾乎所有的構樹都攜帶綠色編號1號的基因單型(</b><b>cp-1)</b><b>或</b><b>cp-1</b><b>衍生的單型,而</b><b>cp-1</b><b>是福建至華中一代最常見的單型</b>。根據古地質孢粉報告與考古證據,帶有cp-1的構樹或許有可能是「出南中國」由福建被帶入北臺灣。但由於現在中國南方已無說南島語系的民族,我們也不知道「南島語系」最早的發源地是在「臺灣」還是「南中國」,因此只能推論<b>北臺灣攜帶cp-1的構樹有可能是「南島語系先祖」由中國南方帶到北臺灣</b>。",
"weight": 65
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"title": "巴布亞新幾內亞地區的基因單型",
"desc": "巴布亞新幾內亞的樣本中,一株帶有cp-17、兩株帶有cp-34。\r\ncp-34 是廣東至中南半島最常見的基因單型,這兩個帶cp-34的樣本或許有可能是源於中南半島。在新幾內亞山區同時發現來自臺灣 (cp-17) 與中南半島的基因單型,似乎與 Triple I 假說的論點所能推論的基因單型分布結果一致。",
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"title": "新幾內亞的構樹",
"desc": "而新幾內亞的構樹似乎也藏著有趣的訊息。<br>新幾內亞的樣本為標本材料,約半世紀前採集自遙遠的山區,3個樣本中,<b>1株帶有cp-17單型、2株帶有cp-34</b><b>單型</b>。cp-34 是廣東至中南半島最常見的基因單型,這兩個帶cp-34的樣本或許有可能是源於中南半島。由於新幾內亞內陸是第一波來到近大洋洲的巴布亞人的居住地,並非南島語族的居住地,在新幾內亞山區同時發現來自臺灣與中南半島的基因單型,似乎與<b>航海廊道三源說</b>的論點十分接近,暗示著新舊往來的人群曾在此處有過充分交流。",
"weight": 66
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"title": "東加_島上栽植的構樹",
"desc": "栽植用於製作樹皮布的構樹,通常會受到悉心照料,除去多餘的側枝,確保樹幹筆直無節,且樹皮完整無孔洞",
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},
"title": "太平洋島上的「構樹田」",
"desc": "雖然從基因單型的遺傳分析可以推論太平洋構樹應該是由臺灣傳入,但要如何證明構樹在太平洋的傳播真的是人類活動所致、而非自然散播呢?<br><br>由構樹在臺灣呈現的顯著遺傳結構我們已知這種植物<b>不太可能靠自身的力量進行長距離的傳播</b>,更別說要橫越廣大的水域,到達一個個的島嶼。<br>而根據文獻,太平洋構樹全賴南島語族栽種繁殖,在18世紀橫渡太平洋的航海日誌中,英國庫克船長 (Captain James Cook) 亦清楚記載,在遠大洋洲島民藉由樹根萌櫱扦插,<b>以無性繁殖的方式栽植構樹</b>。構樹在臺灣隨處可見,常在荒地中恣意生長,但大洋洲的熱帶環境卻讓構樹難以適應,需要人工照顧。研究團隊在太平洋地區進行田野調查時,也發現當地的構樹多是由島民特意栽種,並細心養護。<br>",
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"level": "primary",
"title": "構樹的性別在原生地、近大洋洲與遠大洋洲的分布",
"desc": "採集自各地構樹樣本的雌雄比例,圖中紅色表示雌性,藍色為雄性;圓餅圖的大小與樣本數成正比。可以明顯看出太平洋地區的構樹多為雌性。",
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"title": "請輸入頁面標題",
"desc": "太平洋構樹的主要用途為製作樹皮布,植株在長至3-5公尺時便會被採收,根本來不及成熟繁殖,許多太平洋島民甚至沒有看過構樹開花結果的樣子。且構樹是雌雄異株的植物,兩個性別的個體需要同時存在才能進行有性繁殖,目前的文獻中,幾乎沒有在太平洋地區採集到雄株的紀錄,因此無性繁殖才會成了此區構樹的生殖方式。2016年,<a target=\"_blank\" rel=\"nofollow\" href=\"https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0161148\">智利團隊根據分子標記的分析</a>,證明了太平洋構樹絕大多數都是<b>雌性植株</b>。<br><br>或許,南島語族先祖攜帶構樹旅行時,<b>只帶了雌性植株,之後藉由根部萌櫱無性繁殖的方式</b>,讓構樹生長、傳遍於一個又一個島嶼。<br>",
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"desc": "關於構樹,目前我們已經知道的是<br><b>太平洋構樹起源於臺灣!<br></b><b>太平洋構樹為無性繁殖-</b><b>南島語族引入!<br></b><br>2015年底,美國PNAS期刊刊出了研究團隊的成果—<a target=\"_blank\" rel=\"nofollow\" href=\"http://www.pnas.org/content/112/44/13537\"><b><i>A holistic picture of Austronesian migrations revealed by phylogeography of Pacific paper mulberry</i></b></a>,構樹的DNA不僅記錄了構樹<b>「出南臺灣」</b>遠渡重洋,經蘇拉威西、新幾內亞至遠大洋洲島嶼的歷史,<span>提供支持南島語族<b>「出臺灣說」</b>的有力證據;</span>分佈在福建與臺灣北部的cp-1 似乎也提供了南島語族先祖「<b>出南中國」、「出北臺灣」</b>的線索。而由採集自新幾內亞山區的構樹標本中取得的DNA同時偵測到單倍型cp-17以及中南半島最常見的單倍型cp-34 則顯示了<b>「航海廊道整合說」</b>的可能性。<br><br>而關於構樹的傳播是否能夠推論南島語族遷徙的路徑,我們還需要確定的是<br><span>太平洋構樹是否為古代引入?<br></span>太平洋構樹的傳播路徑是否與人類學推論一致?<br><br>過去10年的努力描繪出了堅實的藍圖,近5年來,人類遺傳學分析古人類基因體也得到了可以相互佐證的結果。然而構樹的研究仍在繼續,研究團隊接下來將一步步聚焦,用更完整的採集取樣填補缺失的足跡、更先進的分子技術增加資料分析的精確,而與更多元的領域結合則能豐富視野,更完整的訴說、呈現人類遷移史上重要的一程。 <b></b>",
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"desc": "<div>5千年前一個適合遠航的日子,風起帆揚,南島語族人整裝待發,船上滿載工具糧食、作物牲畜,要在下一個小島再打造一個家園。他們投入未知的海相,有了星星和海鳥,前方的航道便豁然開朗,漫長的旅程中,他們是否想過,自己走在人類擴張全球版圖的最後一段路。<br><br></div><div>5千年後的今天,來自不同領域、不同國家的學者在同一片海洋及島嶼間奔走,與過往的時空擦肩,這段偉大的海上遷移再次啟航,本研究是第一個以民族植物學切入,以構樹的親緣地理反映了南島語族遷徙的全貌,藉由共生物種親緣地理學證據支持「出臺灣說」,同時也見證了臺灣在南島研究上的重要性。<br><br>●相關展覽<br><a target=\"_blank\" rel=\"nofollow\" href=\"http://brmas.openmuseum.tw/muse/exhibition/23fc9a7b63113167d5e98693ecc46392\">有構厲害|子曰:「必也正名乎。</a><a target=\"_blank\" rel=\"nofollow\" href=\"http://brmas.openmuseum.tw/muse/exhibition/23fc9a7b63113167d5e98693ecc46392\">」</a><br><a target=\"_blank\" rel=\"nofollow\" href=\"http://brmas.openmuseum.tw/muse/exhibition/4c6df6672d9446cad5b03faafe99ed63\">有構厲害|打樹成衣</a><br><br>●聯絡我們<br>中央研究院生物多樣性研究中心<br><a target=\"_blank\" rel=\"nofollow\" href=\"http://www.biodiv.tw/tw/%E4%BA%BA%E5%93%A1/%E7%A0%94%E7%A9%B6%E4%BA%BA%E5%93%A1/%E9%8D%BE%E5%9C%8B%E8%8A%B3\">鍾國芳</a><br>bochung@gate.sinica.edu.tw</div>",
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"fileOversize": false,
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"fileOversize": false,
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"title": "構樹的葉綠體基因組",
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"fileLicence": "著作權利所有",
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"level": "primary",
"title": "構樹葉綠體 ndhF-rpl32 基因單型網狀圖/構樹葉綠體 ndhF-rpl32 基因單倍型網狀圖",
"desc": "各基因單型 (haplotype) 以不同色系表示,並標上編號1-48,跨號中的數字為樣本數,圓圈的大小與樣本數成比例,圓圈越大表示攜有該基因單型的樣本越多,小於10的樣本數未標示。出現在臺灣的基因單型以虛線表示,方形則表示該基因單型為特有基因單型 (unique haplotype)。",
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"fileOversize": false,
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"title": "構樹基因單倍型的地理分布",
"desc": "將不同的基因單型依照分布地區著色;日本:淡橘色,中國:黃色,臺灣:綠色,菲律賓:紫色,印尼:深藍色,近大洋洲:淺藍色,遠大洋洲:橘紅色。\r\n由圖中可以看出,臺灣、中國與中南半島有絕大多數的基因單倍型,為構樹遺傳多樣性中心,極有可能為構樹的原生地。\r\n",
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"fileOversize": false,
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"level": "primary",
"title": "構樹葉綠體 ndhF-rpl32 基因單型分布及網狀圖/構樹葉綠體 ndhF-rpl32 基因單倍型分布及網狀圖",
"desc": "將各基因單型 (haplotype) 以不同色系表示,並標上編號1-48,圖中可見各基因單型於地理上的分布以及網狀圖。\r\n在地圖上以圓餅圖標示不同基因單型於同一地理區所佔的比例,圓餅圖的大小對應該地的樣本數。在網狀圖中各基因單型的編號標示於圓圈內,跨號中的數字為樣本數,圓圈的大小與樣本數成比例,圓圈越大表示攜有該基因單型的樣本越多,小於10的樣本數未標示。出現在臺灣的基因單型以虛線表示,方形則表示該基因單型為特有基因單型 (unique haplotype),僅出現在特定地理區。太平洋地區紅圈內填黑色代表標本樣本。",
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"fileCreator": "鍾國芳",
"rights": "鍾國芳",
"rightsLogo": "http://brmas.openmuseum.tw/files/muse_brmas/styles/square_thumbnail_100/public/default_rights/memder_non-02.png?itok=fGKM5I0r",
"fileLicence": "創用CC 姓名標示-非商業性-相同方式分享",
"fileOversize": false,
"thumb1920": "https://brmas.openmuseum.tw/files/muse_brmas/muse_styles/w1920/mcode/af77ee210df368c42d87a6ff61745314.jpg?itok=0gFKZYLj"
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"title": "福建至華中及臺灣北部地區的基因單型",
"desc": "福建至華中一帶最常見的單倍型 cp-1亦為北臺灣構樹主要攜帶的基因單型。",
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"fileOversize": false,
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"title": "巴布亞新幾內亞地區的基因單型",
"desc": "巴布亞新幾內亞的樣本中,一株帶有cp-17、兩株帶有cp-34。\r\ncp-34 是廣東至中南半島最常見的基因單型,這兩個帶cp-34的樣本或許有可能是源於中南半島。在新幾內亞山區同時發現來自臺灣 (cp-17) 與中南半島的基因單型,似乎與 Triple I 假說的論點所能推論的基因單型分布結果一致。",
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"fileOversize": false,
"thumb1920": "https://brmas.openmuseum.tw/files/muse_brmas/muse_styles/w1920/mcode/5b233385b7756009adce5dca17dd4e0a.jpg?itok=VMcx_SCH"
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"title": "東加_島上栽植的構樹",
"desc": "栽植用於製作樹皮布的構樹,通常會受到悉心照料,除去多餘的側枝,確保樹幹筆直無節,且樹皮完整無孔洞",
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"fileLicence": "創用CC 姓名標示-非商業性-相同方式分享",
"fileOversize": false,
"thumb1920": "https://brmas.openmuseum.tw/files/muse_brmas/muse_styles/w1920/mcode/b63c1c3746baa5fc251816601e144c7e.jpg?itok=1zIJYeoO"
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"title": "構樹的性別在原生地、近大洋洲與遠大洋洲的分布",
"desc": "採集自各地構樹樣本的雌雄比例,圖中紅色表示雌性,藍色為雄性;圓餅圖的大小與樣本數成正比。可以明顯看出太平洋地區的構樹多為雌性。",
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"fileCreator": "Johany Peñailillo,Gabriela Olivares,Ximena Moncada,Claudia Payacán,Chi-Shan Chang,Kuo-Fang Chung,Peter J. Matthews,Andrea Seelenfreund,Daniela Seelenfreund",
"rights": "PLOS one",
"rightsLogo": "http://brmas.openmuseum.tw/files/muse_brmas/styles/square_thumbnail_100/public/default_rights/memder_non-03.png?itok=h0rkcAOw",
"fileLicence": "創用CC 姓名標示-非商業性-相同方式分享",
"fileOversize": false,
"thumb1920": "https://brmas.openmuseum.tw/files/muse_brmas/muse_styles/w1920/mcode/83765eae7ca55f9179cb3286c18931d2.jpg?itok=sVwnSNDs"
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"title": "復活節島田野調查",
"desc": "2014年10月,鍾國芳副研究員於赴智利與合作團隊討論研究計畫,並一同前往野外採集。\r\n研究團隊前往復活節島西南端的知名景點Rano Kau火山口湖,湖壁四周生長的構樹為早期南島語族引入的族群。",
"thumbURI": "https://brmas.openmuseum.tw/files/muse_brmas/muse_styles/square_thumbnail_100/mcode/a3960a2f6bba37646771e9313400a2de.jpg?itok=x2p8VJfU",
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"fileCreator": "鍾國芳",
"rights": "鍾國芳",
"rightsLogo": "http://brmas.openmuseum.tw/files/muse_brmas/styles/square_thumbnail_100/public/default_rights/memder_non-02.png?itok=fGKM5I0r",
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"desc": "特別感謝張至善博士 (C.-S.C.)的研究發想與實驗設計,劉筱蕾小姐 (H.-L.L.)的實驗操作及資料分析。<br><br>本展內容主要參考文獻<br>-劉筱蕾 (2012)<b> 東亞與大洋洲構樹親緣地理學研究 Phylogeography of East Asian and Oceanian Paper Mulberry</b>. 碩士論文. 國立臺灣大學森林環境暨資源學系<br>-Chang, CS, Liu, HL, Moncada, X, Seelenfreund, A, Seelenfreund, D, Chung, KF. 2015. <b>A holistic picture of Austronesian migrations revealed by phylogeography of Pacific paper mulberry</b>. Proceedings of the National Academy of Sciences of the United States of America 112:13537-13542(doi:10.1073/pnas.1503205112) <br><br>●共同研究團隊<br>-國立臺灣史前文化博物館 張至善博士 (C.-S.C.)<br>-智利 Andrea Seelenfreund博士與Daniela Seelenfreund (D.S.)博士研究團隊<br><br>●野外調查協助<br><span>楊親二 (Q.-E. Yang)<br>許為斌 (W.-B. Xu)<br>J. Regalado<br>H. Nguyen<br>I. Kailey (Province Museum of Central Sulawesi)<br>郭弘吉 (H.-C. Kuo)</span><br><br>●研究材料提供<br><span>江友中 (Y.-C. Chiang)<br>彭鏡毅 (C.-I. Peng)<br>劉筱蕾 (H.-L. Liu)<br>K. Nakamura<br>H. Won <br>T. Phuttai <br>楊宗愈 (T.-Y.A. Yang)<br>C. Walter (Pitcairn)<br>B. Kam (Bishop Museum)<br>P.v. Dyke (Amy BH Greenwell Ethnobotanical Garden)<br>M. DeMotta and R. Matthews (National Tropical Botanical Garden, NTBG)<br>K. Aiona (NTBG Kahanu Garden)<br>J. Paman (Maui Nui Garden)<br>B. Mulloy, and D. Tanahy (Hawaii)<br></span><span>Easter Island National Park<br>The herbaria of the Natural Historical Museum<br>New York Botanic Garden<br>Missouri Botanical Garden Smithsonian Institute<br></span>B. Torke<br>V. Funk<br>劉世慧 (S.-H. Liu)<br><br>●研究經費<br>-中央研究院主題研究計畫,整合考古學、生物學、氣象學重建南島語族擴張與遷徙的全史,AS-TP-107-L18<br>-National Science Council, Taiwan Grant NSC-102-2621-B-002-007 (to K.-F.C.)<br>-Fondo Nacional de Ciencia y Tecnología (FONDECYT), Chile Grants 1080633 and 1120175 (to A.S.)<br>-教育部世界南島研究計畫獎助金<br> (to C.-S.C., H.-L.L., and D.S.)<br><br>●參考文獻<br>-張至善(2012)談樹皮布與構樹。林業研究專訊 19(2):62-66 <br>-臧振華 (2012) 再論南島語族的起源與擴散問題。南島研究學報 3(1):87-119<br><span>-Barker, C. (2002) Plate 432.\n<i>Broussonetia papyrifera</i>. Curtis's Bot Magazine 19(1):8–18</span>\n\n<br>-Diamond, J. M. (2000) Taiwan's gift to the world. Nature 403: 709<br>-Gray, R. D., Drummond, A. J., Greenhill, S. J. (2009) Language phylogenies reveal expansion pulses and pauses in Pacific\nsettlement. Science 323(5913):479–483\n\n<br>-Green, R. C. (2000) Lapita and the\ncultural model for intrusion, integration and innovation. Australian\nArchaeologist: Collected Papers in Honour of Jim Allen, eds Anderson A, Murray\nT (Coombs Academic Publishing, Australian National University, Canberra,\nAustralia), pp 372–392\n\n<br>-Herb Kawainui Kane (1998) Ancient\nHawaii. Kawainui Press\n\nKo AM-S, <i>et al</i>. (2014) Early Austronesians:\nInto and out of Taiwan. Am J Hum Genet 94(3):426–436\n\n<br>-Lipson, M. (2018) Population Turnover in Remote Oceania Shortly after Initial Settlement. Current Biology 28:1157–1166 <br>-Oppenheimer, S. J., Richard, M. (2001) Polynesian origins. Slow boat to Melanesia? Nature 410(6825):166–167 <br>-Matisoo-Smith, E., Robins, J. H. (2004)\nOrigins and dispersals of Pacific peoples: Evidence from mtDNA phylogenies of\nthe Pacific rat. Proc Natl Acad Sci USA 101(24):9167–9172\n\n<br>-Matisoo-Smith, E. (2014) Ancient DNA\nand the human settlement of the Pacific: A review. Journal of Human Evolution 79:93-104\n<br><span>-Peñailillo, J., Olivares, G., Moncada,\nX., Payacán, C., Chang, C.-S., Chung, K.-F., Matthews, P. J., Seelenfreund, A.,\nSeelenfreund, D.. 2016. Sex distribution of paper mulberry (<i>Broussonetia\npapyrifera</i>) in the Pacific. PLoS One 11:e0161148\n(doi:10.1371/journal.pone.0161148)</span>\n\n<br>-Posth, C., <i>et al</i>. (2018) Language continuity despite population replacement in Remote Oceania. Nature Ecology & Evolution 2:731–740 <br>-Skoglund, <span>P., <i>et al</i>. (2016) Genomic\ninsights into the peopling of the Southwest Pacific. Nature 538:510–513</span>\n\n<br>-Nielsen, R. , Akey, J. M. , Jakobsson, M. , Pritchard, J. K. , Tishkoff, S. & Willerslev, E. (2017) Tracing the peopling of the world through\ngenomics. Nature 541:302–310<br>-<span>Seelenfreund, D., <i>et al</i>. (2010) Paper\nmulberry (<i>Broussonetia papyrifera</i>) as a commensal model for human mobility in\nOceania: Anthropological, botanical and genetic considerations. NZ J Bot\n48(3-4):231–247</span> <br><br>●數位典藏與數位人文學研究計畫 AS-ASCDC-107-304 <br>",
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來源
中央研究院生物多樣性研究中心 (BRCAS)